My grandfather was a lifelong Republican but also a great lover of the outdoors and the national and state park systems. The idea of putting up golf courses on park land would have sickened him. He wanted people in the future to have the same opportunities he had to enjoy camping, hiking, etc.
I don’t know why Rubio and Mast chose this moment to publicly fall out with DeSantis, but among the rank-and-file Republican voters there have always been individuals who cared about conservation.